Wallacea: A Biodiversity Hotspot
Overview
Vertebrate diversity
Plant diversity
Overview

Wallacea

The Wallacea region is located almost entirely within the borders of Indonesia and includes the large island of Sulawesi, the Moluccas (Spice islands), Banda islands and the Lesser Sundas. The hotspot covers a total land area of total land area of 338,494 km².

The Lesser Sundas are located south of Sulawesi, and include Bali, Lombok, Sumba, Sumbawa, Flores and Timor. The Moluccas includes several hundred islands in the north-east of the hotspot, the largest being Seram and Halmahera.

Wallace's line

The Wallacea hotspot is separated from the adjacent Sundaland hotspot to the west by Wallace's line, which runs between Borneo and Sulawesi to the north and Bali and Lombok to the south. Wallace's line is named after the British naturalist Alfred Wallace, who noted distinct bird and mammal faunas on either side of the line. Wallacea is a transition zone between Asian and Australasian regions, containing flora and fauna from both of these areas. Many mammals originate from Asia whilst birds have Australasian origins.

Why is Wallacea high in endemic species?

Wallacea contains a high number of endemic species. Several factors explain why the region in rich in endemics:

1.) Tropical climate: High temperatures are thought to lead to high rates of mutation, and therefore high rates of speciation.

2.) The region contains many islands: Isolation is a major factor leading to the formation of endemic species. Islands are isolated because they are surrounded by areas of sea, which represents a barrier to migration. Isolated species on islands may evolve into unique species adapted to their local habitat/ecological niches.

3.) Geological history: Wallace's line marks the position of deep ocean trenches and separates shallow waters to the west from the deeper waters to the east. During past ice-ages, sea levels dropped and the shallow waters to the west disappeared, allowing migration of fauna between the islands of Java, Borneo, Sumatra and Bali and therefore spread of similar species between them. Islands to the east in the Wallacea region remained isolated by the deep ocean waters, preventing migration of species and leading to the evolution of distinct faunas.

Wallace's line

Although Wallacea contains fewer species overall compared to the Sundaland hotspot to the west, the region contains a very high number of endemic species for it's size:

Table 1: Total number of species and number of endemic species for various taxonomic groups

Taxonomic group
Number of species
Number of endemic Species
Percent endemism
Plants
10 000
1500
15
Mammals
222
127
57.2
Birds
647
262
40.5
Reptiles
222
99
44.6
Amphibians
48
33
68.8
Freshwater fish
250
50
20

 

 

 

 

 

Ecosystems

The principle terrestrial vegetation is tropical rainforest, but there are also large areas of savannah grassland on the Lesser Sundas. The Wallacea hotspot also includes some of the most biologically diverse coral reefs in the world. The World Wildlife Fund lists several regions in Wallacea as Global 200 ecoregions, a collection of the Earth's most biologically diverse and representative terrestrial, freshwater, and marine habitats. These include Sulawesi moist forests, Moluccas moist forests, Nusa Tengarra (Spice islands) dry forests, central Sulawesi lakes and the Sulu-Sulawesi and Banda-flores seas (tropical coral).