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| Swabian plesiosaurs and Tanzanian dinosaurs (as viewed
from 1909)
The following is my translation of an article
called: Paläontologische Umschau (Schwäbische Plesiosaurier und ostafrikanische Dinosaurier)
nach Vorträgen von Prof. Dr. E Fraas, zusammengestellt von Dr. K Floericke. It appeared in a German
popular science magazine, Kosmos Handweiser für Naturfreunde 1909, Heft 1, Seiten
1-5. Some of the information is naturally out of date. I'm not aware of any
previous translation.
Paleontological review (Swabian plesiosaurs and East African dinosaurs) according to
reports of Prof Dr E Fraas, presented by Dr K Floericke
The reptile world of the Swabian Jurassic Formation has long been known for the
excellent completeness and beauty of Ichthyosaurus, better specimens of which could
hardly be hoped for. The preservation of these pre-Diluvium beings is sometimes so good,
that not only can one see the precise appearance of their bodies, one can also reach
conclusions about their lifestyle. We find ichthyosaurs with skin impressions and
also females carrying either their own young within their bodies or perhaps those who
had been swallowed. And the number of available specimens is now so high that jokingly,
but not inappropriately, the Ichthyosaurus has been named 'the Swabian
pet'.
In contrast, remains of the related plesiosaurs have always been very rare, and only
consisted of isolated bones from spines or feet until, in the year of 1893, the
famous preparator, Bernhard Huff, had the luck of bringing an almost complete skeleton
to light from Holzmaden but, due to a lack of funds, this went to Berlin rather than
staying in the area, and there it was more closely researched by a geologist, Dames,
and described as Plesiosaurus guilelmi imperatoris. As Professor Fraas emotionally
put it in one of his recent addresses, upon which these lines have been based, he can
never go past this glorious specimen in the Berlin Museum without closing his eyes, as
his loss is so heartfelt that this treasure from the ancient Swabian Jurassic sea is not
housed in Stuttgart. In a remarkable and unexpected way, this wish has been fulfilled
in the past year as, not only did the Wüttemberg collection gain plenty of new
material from the Triassic of Swabia, the Jurassic of Africa and the Tertiary of
Egypt, but it also simultaneously received two plesiosaurs from Holzmaden, and these
are exceptionally fine and well preserved. Scientifically speaking, the interest
with this is that both these valuable items, which have recently provided a fine
exhibit for the Stuttgart Naturalienkabinett, involve two separate species of
the widespread family of plesiosaurs, each apparently from the most extreme grades.
No fewer than 26 species have been reported from the English Liassic, although many of
these are based on poor and difficult to diagnose remnants, but all seem to fit
comfortably between both Swabian examples. One of this pair lay in the so called
Stinkstein, a very hard stratum from the upper Liassic shales, and this made
freeing it from the steel hard matrix especially difficult. Normally, one does not
attempt to prepare finds such as this from the rock as the effort is not worthwhile,
and the fragile bones break. However, with such an exceptional find, there could naturally
not be such an uncourageous reluctance, and the skill of Hauff actually succeeded,
after nine months of arduous work, in bringing this Plesiosaurus to sight
and, in such an excellent condition, it can be termed the best preserved
Plesiosaurus in the world. It was the stomach side which was exposed and, in
line with experience, that is both best suited for withstanding such efforts and, in
the case of Plesiosaurus it also provides the diagnostic details. The
circumstances were much more favourable with the second specimen from the Liassic shale,
as it was embedded in much softer rock. This second example is shown on its side in
what looks like a swimming position, and it belongs to the same species as the Berlin
specimen excepting for it being significantly larger with a body length of 340cm,
while the Berlin one measures 280cm. However, as the mode of the bones and various
other features clearly indicate we are dealing here with a younger animal when it
comes to the Berlin Plesiosaurus and a full grown adult with the Stuttgart
specimen, we may assume that, despite the differential in size, both cases belong to
the same species, that is Plesiosaurus guihelmi imperatoris.
The first found and arduously salvaged specimen from the Stinkstein looks
significantly different. While it is only slightly longer (344cm), it nevertheless
appears to be much more massive, and the build of its back make it look more gigantic.
It has been named Plesiosaurus victor in honour of Herr Victor Fraas, the
brother of Professor Fraas, who provided much of the money required to purchase this
fine piece. While Ples. gui. imp. is a smaller and graceful creature, due
namely to its long neck and small head, Ples. victor had a much plumper,
heavier, so to say a pre-Diluvium appearance.
The most obvious features of the stomach region are the large, plate-like bones of
the compressed breast area, which correspond to the also plate-like developed hips,
and the wondrous confusion of false ribs between the two, so called stomach ribs
similar to those of fish, a basket-like structure. This was obviously for protection
against collisions with stormy sea waves. The structure of the extremities also
speaks for a lengthy stay in the sea as they have developed into proper paddles, and
obviously served as rowing organs, while the ribbed tail had the function of a
rudder. In some respects the skull is reminiscent of a lizard, but it is proportionately
very small and flat and armed with sharp teeth, which sat in their alveolae as with
crocodiles of the present day. The nature of this dentition shows that plesiosaurs
were not plant-eaters but rather predators, and they must have mainly lived off fish
and squids from the Jurassic sea. The neck of these remarkable animals was uncommonly
long, consisting of between 24-41 vertebrae and yet it would have been relatively
immobile. We should not imagine that Plesiosaurus could perform snake-like
movements like those of a swan. That would already be impossible due to the character
of the neck vertebrae which, excepting for the first two, have short ribs on their
keels as the neck bones become very thin due to hollows, and these allow them to fit
tightly against each other as if they were flat plates. The long neck was especially
suited for fetching all kinds of small animals from out of the soft floor in coastal
areas, or for sudden whip-like attacks on prey. It may also be that the length of
the neck allowed the animal to keep its head above the breaking waves. As Professor
Fraas reported, the life and activities of the African Schlangenhalsvögel ('darters'),
snake-necked birds of the great African lakes, provide us with a living and visible
image perhaps reminiscent of plesiosaurs, when it comes to how these fabled animals
might have moved and dived in the Jurassic sea of ancient Swabia.
It is interesting to compare plesiosaurs with the contemporary, but very distantly
related ichthyosaurs, as this shows the creative energy of nature can often simultaneously
use very different methods to reach the self same objectives. Both animals evolved
from land dwellers and had to adapt to being aquatic, and this end was reached by
entirely different routes. With the Ichthyosaurus, which externally compares
with our extant dolphins, we see the most magnificent principle of a torpedo, whereas
Plesiosaurus is more like a flat boat. If the first may have shot through the
waves like an arrow, then the latter appears slower and more massive, more suited to
resisting the crashing waves of the coastal zone so as to move in that area. And
just as the basic build of Ichthyosaurus is still maintained by today's
dolphin, so the present also provides a refuge for an animal whose construction strongly
reminds us of the Plesiosaurus, namely the marine turtle. Protection against the
power of the waves is provided by the rib cage and stomach ribs of Plesiosaurus,
whereas the marine turtle achieves the same with the armour on its underside.
Although science has not yet managed to identify stem-forms for the
Ichthyosaurus among terrestrial animals, and make the still silent branches
accessible for natural historians, more progress has been possible in this regard
for the Plesiosaurus; we do know animals from the Muschelkalk Formation
which obviously stand in a close relationship to the plesiosaur, namely the nothosaurs,
creatures which may have lived predominantly on land, but which already showed strong
adaptations for aquatic conditions. And small, completely terrestrial nosticosaurs,
found in Lettenhöhle Cave near Hornegg, which may have been pygmies in comparison
to the Plesiosaurus with their lengths of 25-30cm, but nevertheless seem
anatomically and morphologically directly connected with nothosaurs. We are also here
in the fortunate position of being able to follow a transition from terrestrial lifestyles
to aquatic animals in various developmental stages; the phenomenon, that aquatic
residents in the endless ocean develop themselves into true giants when compared to
their ancestors, is rather too common in nature to be accepted as something that is
exceptional. It is to be expected that the Jurassic of Swabia will provide more
highly interesting fossils in the future, and that not only the fantasy, but strict
science will appreciate an ever richer haul of remarkable animal figures lived in the
Jurassic sea of Swabia.
During the Jurassic and Cretaceous periods, we find a vast development of the reptile
world not just in the ocean, but also on the main land that was populated by reptiles
of a size and form which is unique. Prime among these are the 'terrible lizards' or
dinosaurs with their fantastic forms and, sometimes, truly gigantic sizes. The Upper
Jurassic of Wyoming has provided Diplodocus, known from a complete skeleton*
of 22m in length, while bones and skeletal remains of Brontosaurus indicate
an animal of over 30m long. As well as the widely known discovery sites of England
and North America, we now have a new and important locality in the south of our East
African province, which has been discovered and investigated by Prof. Fraas in
recent years.
(* The wonderfully produced replica of this skeleton, a present for Kaiser Wilhelm
from Carnegie, is displayed in the Lichthof of the Museum of Naturkunde
in Berlin, and the Senckenberg Museum in Frankfurt also possesses a large part of a
skeleton.)
As this academic recently reported in the Berlin Gesellschaft für Erdkunde,
he discovered a great quantity of prehistoric giant animals on the Mbemkuru River
but, unfortunately, at a time when his health was weakened by dysentery and he was
otherwise insufficiently equipped for a large expedition. Furthermore, all ground in
the area was covered by a healthy growth of bush, and could only be made accessible in
places when cleared by burning; but that made it apparent that parts were so rich with
such piles of bones, that this locality is on a par with the famous locations in
North America. Although only cursory investigations were possible, there can be no
doubt that this newly discovered Fundstätte promises science a very rewarding
booty over the forthcoming decades, and some of these fable creatures will be
previously unknown or only poorly known, thus providing new information on prehistory,
but only when it is possible to systematically and thoroughly investigate and secure
the remains, and to bring them to European museums. In order to aid an understanding
about these giant bones, it would be best to state that, for example, a single upper
thigh bone presently on display in Stuttgart reaches a length of 1.4m and a mass of
three hundredweight, and there are vertebrae as large as those from the giant
Diplodocus and Brontosaurus. The individual bones of this Upper
Cretaceous formation, taken as a whole, are reminiscent of North American localities
and are very well preserved although, unfortunately, finds from the surface have
been badly damaged by weathering, and that makes searching for good skeletons sometimes
very difficult. As Fraas assumed that still complete skeletons could be found in at
least deeper layers of the ground, he carried out an excavation on a promising looking
slope, and a partial articulated skeleton was soon discovered, part of which was
delivered to Stuttgart. A scientific examination concluded this belonged to a new
group of huge dinosaurs which were named Gigantosaurus. Fraas differentiates
between a larger form, G. africanus, and a more compact one, G.
robustus.
Additional Note
An index of more of my translations of old Kosmos articles can be found at:
A number of Mesozoic (and post-Mesozoic) location summaries can be found at
Localities.
http://home.arcor.de/ktdykes/meseucaz.htm |